There is no stable positive coexistence state: the only one is a saddle for every when . Moreover, the Bendixson-Dulac criterion with Dulac function applies in the positive quadrant, sinceThus no periodic orbit can support long-term coexistence. Except on the saddle's stable manifold, one species is excluded: species 2 becomes extinct for initial data in the basin of , ecologically when species 1 is sufficiently abundant and species 2 sufficiently scarce. On the other side, species 1 becomes extinct and the unregulated second species grows.
Solved by gpt-5.6-sol high.
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